what is optimum temperature for photosynthesis

The Four Substances Recycled In Photosynthesis And Respiration, The Potential Dangers Of Overdosing On Neem: What You Need To Know. 114, 439-444. The Optimum Temperature For C4 Plant Growth [Source: Authors diagram]Differences in the thermal response of photosynthesis are also found in individuals of the same species growing at different temperatures. There are three major temperature points where a plant grows. Expand. What Is The Optimum Temperature For Photosynthesis And Why? Consider the temperature. Acquired tolerance to temperature extremes. During the winter growing season, photosynthesis is stressed, resulting in changes in photosystem II electron transport. Three Elodea will be placed into a beaker containing a diluted potassium bicarbonate solution and placed on ice. The rate decreases with an optimal temperature (usually around 35 degrees Celsius). The same reasons as above may explain the decrease in CO2assimilation when the temperature increases beyond that of the thermal optimum. Why would it be hard to find the ideal light intensity if the temperature were very hot or cold. However, at high temperatures, enzymes are denatured and this will decrease the rate of photosynthesis. Remth (Hammada scoparia), a characteristic plant of the Wadi Rum desert (Jordan). The enzymes that are responsible for the photosynthetic reaction are not affected by changes in temperature. Also, the amount of light that's available is important. Photosynthetic adjustment to temperature. Are temperature sensitive: Figure 9. around 37 oC What is the optimum temperature for photosynthesis and why? One can then wonder about the interest of this change. This water allows, among other things, the cooling of the leaf (see Focus Leaf transpiration and heat protection). The most distinctive feature is a shift in the optimum temperature of photosynthesis as the growth temperature changes and enabling them to maintain photosynthetic efficiency under such environment (Yamori et al., 2014). It highlights the temperature range in which variations are generally rapidly reversible. At least, that is, up until 25 degrees Celsius. At low temperatures, the rate of photosynthesis is limited by the number of molecular collisions between enzymes and substrates. ATP is synthesized when protons from the lumen return to the stroma through an ATPase using inorganic phosphate, Pi. Some crops, such as cane and maize, prefer warm temperatures to germinate. Sci. There is no way to quantify the extent to which Elodea has an effect on change in the amount of water it is found in. When photosynthesis is limited by high temperatures, it has a limited rate of rate due to the number of molecular collisions between enzymes and substrates. Optimal temperatures for GROSS photosynthesis in plants are 15-40C, but above ~10, dark respiration goes up and NET photosynthesis goes down (Fig.). Figure 14. The open circles in subpanel (b) were not included in regression. Ribulose 1,5-bisphosphate carboxylase/oxygenase is a CO2-O2 enzyme. When the temperature rises, the particles in the reaction move quicker and collide more, so the rate of photosynthesis rises also. Terrestrial plants are regularly subjected to strong temperature variations. Begin typing your search term above and press enter to search. Plants use some of the oxygen. photosynthesis is carried out in our bodies. The Power Of Neem DE: A Natural And Effective Insecticide, Getting Rid Of Pesky Caterpillars With Neem Oil, Understanding The Benefits And Risks Of Giving Neem To Your Dog. At higher temperatures, photosynthetic rates may begin to decline due to heat stress on the plant. How does temp affect photosynthesis GCSE? The optimum temperature range is 25 to 30 degree celsius. Figure 7 illustrates the case ofAtripex lentiformis, [7] a perennial leafy plant, which occurs in California in both Death Valley and in cool, wet coastal habitats: Figure 8. [9] Yamasaki T, Yamakawa T, Yamane Y, koike H, Satoh K & Katoh S. (2002) Temperature acclimation of photosynthesis and related changes in photosystem II electron transport in winter wheat. Rubisco activase constrains the photosynthetic potential of leaves at high temperature and CO2. When temperature is increased from minimum to optimum, the rate of photosynthesis doubles for every 10C rise in temperature. Jean-Franois MOROT-GAUDRY, INRA Emeritus Research Director, the Jean-Pierre Bourgin Institute, INRA Versailles, Member of the Acadmie dAgriculture. These cookies help provide information on metrics the number of visitors, bounce rate, traffic source, etc. As a result, it is critical that photosynthesis be stimulated by light in the blue and red ranges. Hypothetical reactions are characterized by the breakdown of the bodys organs and tissues. As the temperature is increased enzyme activity increases to a maximum value at the optimum temperature (around 37 oC for most human enzymes). The temperature-dependence of photosynthesis and respiration was fitted to empirical models to obtain maximum metabolic rates and thermal optima. [Source: Authors diagram]Below 10C and above 34C plants start to set up protective mechanisms. For example, enzymes in the small intestine have an optimum pH of about 7.5, but stomach enzymes have an optimum pH of about 2. In an O2-poor atmosphere(Figure 10), competition between O2and CO2is very reduced. In high temperatures, however, enzymes become denatured, which decreases the rate of photosynthesis. CO2enters the Benson-Calvin cycle and the photosynthetic fixation of O2 is at the origin of a metabolic pathway responsible for photorespiration (Figure 11; see also The path of carbon in photosynthesis). They mainly colonize cool and humid environments (or seasons). Plants become more resistant to otherwise fatal temperatures by being exposed to cool or high temperatures. Various environmental parameters influence the thermal optimum of C3 photosynthesis. Cane and maize, for example, can be used as examples. Plant Physiol. The optimum temperature for most plants is 15C to around 40C. Because of the higher temperatures, proteins in enzymes degrade more quickly. The most important factor in plant growth is light because it aids in photosynthesis and photosynthesis. There is no definitive answer to this question as it depends on the specific plant species involved. In fact, optimal photosynthesis rates typically occur when temperatures are between 77-86 degrees Fahrenheit. The purpose of this study was to determine the effect of light intensity on pondweed photosynthesis. The rate of photosynthesis decreases when the temperature reaches 40 C. It was discovered in 1905 by F.F. Figure 15. Light near saturation. they produce, but most of it is released. It is not a given; the plant in question is a different kettle of fish. It is estimated by the saturation deficit of the partial pressure of water vapor in the ambient air around the leaves. Photosynthesis is the process in green plants by which they create carbohydrates (and the sugar glucose) by absorbing carbon dioxide from the air and water from the soil. Plants in warm environments, or growing during the warm season, have a higher photosynthesis at high temperatures. Short-day potentiation of low temperature-induced gene expression of a C-repeat-binding factor-controlled gene during cold acclimation in Silver Birch. The range shown in Figure 14B does not have an effect on electron transfer in the thylakoid membrane. Simple experiments carried out by scientists shows that the rate of photosynthesis is critically dependent upon variables such as temperature, pH and intensity of light . Photosynthesis is a process that produces organic molecules from simple inorganic molecules from the suns energy. The Elodea will be placed in a beaker containing a diluted potassium bicarbonate solution and placed on wet ice after being diluted. Home Plants will show increased resistance to other types of heat at higher temperatures if they are kept at a cool or high temperature. Desert plants require the greatest amount of light and require a very small amount of water in the soil and air. [Source: Ji-Elle, CC BY-SA 4.0, via Wikimedia Commons]For example, the photosynthesis of Hammada scoparia, a bush in the deserts of the Middle East (Negev, Wadi Rum) follows the seasonal variations in temperature: its thermal optimum varies from 29C in early spring to 41C in summer and then to 28C in autumn. Optimum temperature for seed germination 25oC to 35oC. What are the diverse types of responses? If the persons temperature keeps dropping, they should seek medical attention. Trends Plant Sci. Temperature, carbon dioxide concentration and light intensity can affect the rate of photosynthesis. [Source: Lomvi2, CC BY-SA 3.0, via Wikimedia Commons]For example, the thermal optimum for CO2assimilation [4] in Deschampsia antarctica (Figure 3) and Colobanthus quitensis, the only two Antarctic flowering plants, is between 8 and 15C, while it is around 45C in Tridestomia oblongifolia, a warm desert plant from Central America. So, in general, temperature affects photosynthesis by increasing the rate of the reaction up to a certain point. [Source: Authors diagram]The rapidity of current climate change makes it necessary to delve deeper into the responses of plants to their environment: the hope is to be able to maintain sufficient primary production to keep the biosphere functioning. The enzymes that convert sunlight into energy perform poorly at temperatures below freezing. It is a commonly held belief that high temperatures can adversely affect the rate of photosynthesis in plants. When discussing photosynthesis, it is important to understand that there is an optimum temperature for this process to occur. Does Neem Oil Stain? One of the most important components of photosynthesis is light. Plant. Temperature also plays a part. Diagram of the interacting processes during photosynthetic CO2 fixation (case of a C3 plant). Which is correct poinsettia or poinsettia? The optimum temperature for most plants is 15C to around 40C. If the temperature rises above a certain level, it is generally more common for photosynthesis to occur, and the rate of chemical reactions increases as the temperature rises. Diagram of the variation of CO2 assimilation by an intact leaf. The brain and the body are unable to function properly when the body temperature is dangerously low. The rate of photosynthesis increases with increasing temperature. Human activities lead to an increase in carbon dioxide in the atmosphere. It is thus possible for their growth to be hampered or for them to perish in such a drastic manner. The lumen protons have two origins: (1) oxidation of water in the lumen by PSII which also provides electrons, e- and (2) operation of a proton pump in the thylakoid that passes protons from the stroma into the lumen. How does photosynthesis react to temperature fluctuations? Higher CO2 and N levels have an effect on photosynthetic thermotolerance in plants, which is critical in predicting how plants will respond to global warming. Temperature affects the rate of photosynthesis in crop plants and affects where certain crops can be grown. Limiting factors. The suns energy is converted into chemical energy that is stored in the bonds of the organic molecules. This cookie is set by GDPR Cookie Consent plugin. In normal air, the effect of O2 on photosynthetic CO2fixation (Figure 11) is very low (or even nil) when the temperature is low: competition on the carboxylation sites is in favour of CO2. 3 What is the optimal conditions for photosynthesis? (Yellow flowers) is a typical plant of dry areas in California (here Palm Canyon trail). As a result, higher temperatures typically increase the rate of photosynthesis, making it a chemical reaction, and the rate of most chemical reactions also increases. There is a thermal optimum, similar to that found in plants C3 and C4, for the electron transfer chain. eHow may earn compensation through affiliate links in this story. Carbon dioxide concentration must be taken into account as well. Some plant seeds, including cool season vegetables like lettuce and broccoli, germinate best in temperatures between 55 and 70 degrees F. (13-21 C.), while warm season plants such as squash and marigolds, germinate best when temperatures are between 70 and 85 degrees F. To perform photosynthesis, plants need three things: carbon dioxide, water, and sunlight. The Power Of Neem DE: A Natural And Effective Insecticide, Getting Rid Of Pesky Caterpillars With Neem Oil, Understanding The Benefits And Risks Of Giving Neem To Your Dog. Plant Physiol. A sunny path covered by a canopy of trees. Growers who use indoor grow lights should be aware of these factors in order to keep their plants healthy. For these extreme values,CO2assimilation is often unstable and can be cancelled more or less quickly: the leaf is then under stress (Figure 2). Note that hardening allows leaf maintenance in perennial leaf plants and therefore minimizes energy loss under extreme temperature conditions. The t test will be used to determine the significance of each temperature data set. This cookie is set by GDPR Cookie Consent plugin. Light with a shorter wavelength has a greater impact on photosynthesis than light with a longer wavelength. is as high as 3%. A greenhouse or garden that receives adequate light and water but becomes too hot, resulting in less vigorously produced plants. Due to the increasing frequency of climate change, it is critical to better understand the plants responses to the environment. Producing CO2 by pumping gas into your greenhouse or indoor garden can help ensure a healthy CO2 supply if you have a densely packed greenhouse. Does Neem Oil Stain? Figure 13. The assimilation of the desert ecotype (Figure 7A) and the coastal ecotype (Figure 7B) show almost the same response to temperature when grown under 23C during the day and 18C at night (in red in Figure 7). Ehleringer JR., Sage RF, Flanagan LB, and Pearcy FW (1991) Climate change and the evolution of C4 photosynthesis. To cite this article: CORNIC Gabriel (September 27, 2021), Effects of temperature on photosynthesis, Encyclopedia of the Environment, Accessed March 4, 2023 [online ISSN 2555-0950] url : https://www.encyclopedie-environnement.org/en/life/effects-temperature-on-photosynthesis/. The ideal temperature for photosynthesis is between 70-85 degrees Fahrenheit. Environmental Science. CO2 content in ambient air: 390 ppm. . Figure 12. Oxygen production increases from 10 to 30 degrees Celsius to 30 to 40 degrees Celsius, but decreases from 30 to 40 degrees Celsius to 10 degrees Celsius. Temperature has an impact on the rate at which photosynthesis takes place. The optimum temperature of photosynthesis is the temperature at which the rate of photosynthesis is the fastest. Plant Physiol. How does photosynthesis reacts to rapid and slow temperature variations? Figure 5. This disrupts the plants ability to absorb nutrients from the surrounding soil, which plants secrete enzymes to digest. Carbon dioxide levels and temperatures also have an impact on photosynthesis in a potentially dramatic way. Other temperatures such as 68 and higher, the enzymes start to work less efficiently and above 104 is where the enzymes start to lose their functionality and shape. In general, the photosynthetic process refers to the transfer of energy from an object (usually the sun) to the plant. However, in general, the optimum temperature for photosynthesis is between 20-25 degrees Celsius. To date, most models use the biome-specific optimum air temperature parameter. Maintaining plants at cool or high temperatures causes, along with the changes in photosynthesis described above, increase in their resistance to otherwise lethal temperatures(frost and high temperature). How is temperature changed during photosynthesis? The optimum temperature for photosynthesis is between 20 and 30 degrees Celsius. [Source : Royalty-free image / Pixabay]In a world with steadily increasing atmospheric CO2(Figure 13), the thermal optimum of C3 plants is expected to increase. Jordan DB is one of the authors of the novel Ogren WL. Geophysical Research Letters. At temperatures below freezing, photosynthesis fails to function properly. Higher temperatures increase the number of collisions that plants will have with one another, resulting in an increase in photosynthesis. When temperatures are too hot or too cold, the body produces less oxygen; however, when temperatures are too hot or too cold, the body produces more oxygen. The reason for this is that enzymes that are responsible for the chemical reaction of photosynthesis are more active at higher temperatures. Temperatures, pH, light intensity, and other factors play a significant role in determining the rate of photosynthesis. The assimilation of CO2 by plants via photosynthesis is the gateway to carbon in the biosphere. There are, however, some caveats to this general trend. Climate Change 2007: The Physical Science Basis. Photosynthesis Res. They are included in the thylakoid membrane, which is made up of two lipid layers forming sacs in the chloroplast. The CO2/O2 specificity of ribulose 1,5-bisphosphate carboxylase/oxygenase. Photosynthetic CO2 uptake varies with temperature. Seasonal variations in temperature determine the amount of carbon dioxide in the air, which is what photosynthesis of Hammada scoparia occurs in the Middle East. The Encyclopedia of the Environment by the Association des Encyclopdies de l'Environnement et de l'nergie (www.a3e.fr), contractually linked to the University of Grenoble Alpes and Grenoble INP, and sponsored by the French Academy of Sciences. [6] Ecotype: Plants of the same species from different environments, which, grown from seed to flower under identical conditions show different physiological characteristics. As a result of temperature changes, the rate at which photosynthesis takes place is affected. Light takes 24 hours to penetrate plants during photosynthesis. enzyme denatured at high temperatures. Increased temperatures have not only destroyed the enzymes that carry out photosynthesis, but they have also disrupted the entire photosynthetic process. [7] It fetches water from as far as the water table, hence its name of phreatophyte plant. A variety of crops ability to withstand both heat and cold varies. Green plants and algae use light energy to make glucose and oxygen from carbon dioxide and water. However, there are organizational differences between these two types of plants (see The path of carbon in photosynthesis). The rate at which photosynthesis takes place must be highly influenced by factors such as temperature, pH, and light intensity. It is located around 30C and corresponds to that of CO2 assimilation when the latter is saturating [9]. How Long After Using Neem Oil Should You Wait To Harvest Your Crop? At temperatures greater than 104 degrees Fahrenheit (40 degrees Celsius), photosynthesis degrades its shape and function, causing the rate of photosynthesis to rapidly decrease. C3 plants accounted for approximately 85% of current plant species in the first generation. may not act antagonistically with UVB to mitigate its negative effects when the temperature exceeds the organisms' optimal temperature. Out of these, the cookies that are categorized as necessary are stored on your browser as they are essential for the working of basic functionalities of the website. As a result, low temperatures can have a significant impact on plant growth and health. In low-light, ferns and moss thrive, as their soil is moist and their humidity is high. However, other factors are at play since CO2 assimilation measured (1) in an atmosphere with little or no photorespiration (ambient O2 content of 1%), and (2) measured in a normal atmosphere in a C4 plant decreases in both cases (Figure 10). When the concentration of carbon dioxide in the air is high, photosynthesis is more efficient. Do photosynthesis continues to increase with temperature? At the optimum temperature, the enzyme is most efficient and the rate if maximum. The optimal temperature for photosynthesis varies depending on the type of plant. In this range of temperatures it presents a maximum value: a thermal optimum. Encelia sp. European community. A greenhouse or garden that is too hot, too cold, or too light-producing to produce as much as it should. Because the carbon dioxide concentration increases with the increase in temperature, the heat required to remove it also increases, resulting in a trade-off. (C6H12O6) from carbon dioxide (CO2), and water (H2O). What is the thermal amplitude that allows it to function? 6 How does temp affect photosynthesis GCSE? Above this temperature the enzyme structure begins to break down (denature) since at higher temperatures intra- and intermolecular bonds are broken as the enzyme molecules gain even more kinetic energy. Plants use more photosynthesis during cold growing seasons because the temperature of the environment is lower. Generally speaking, photosynthesis of a plant will increase with an increase in temperature. The optimum temperature for photosynthesis is about 29 C in tropical rainforest (Liu 2020), with a drop in photosynthesis at supraoptimal temperatures (Slot and Winter 2016), which may. The optimal temperature for photosynthesis in the leaves of the 15C, 25C, and 35C ranges was 15C to 20C, 25C to 30C, and around 35C (Fig. Plants begin to develop defenses at temperatures below 10C and up to 34C. How Plants Sense and Respond to Stressful Environments. The respiration of the plant will increase, though, which means it will need more carbon dioxide and water to maintain itself at higher temperatures, even though it will only be growing at its maximum rate. The main variables which affect photosynthesis are light, water, CO 2 concentration and temperature. The reason for this is that enzymes are temperature sensitive and can be destroyed at higher temperatures. Figure 3. When the temperature rises, I believe the rate of photosynthesis will also rise until the temperature reaches 40 Celsius. O2is mainly fixed if its content increases or if that of CO2decreases (the latter then releases active sites which are then occupied by O2). In Sanchez Dias M, Irigoyen JJ, Aguirreolea J & Pithan K (eds) Crop development for cool and wet regions of Europe. By Hardening perennial leaf plants, you can manage leaf maintenance while lowering energy loss at high temperatures. If photosynthesis is to happen at its maximum rate, all environmental factors . In the graph above, as the pH increases so does the rate of enzyme activity. CO2is concentrated at the Rubisco by a mechanism that is insensitive to oxygen. #Heart disease The cell anemia. At a temperature of 8C or higher, CO2 assimilation is best performed. However, anyone can become affected by hypothermia, with the most common cause of damage being an elderly person, a history of medical problems, or a person who is physically inactive. 115, 479-493, Berry J & Bjrkman O (1980). Serge PLANTON, General engineer of bridges, waters and forests, former climatologist researcher at Mto-France, CNRM (Centre National de Recherches Mtorologiques). Temperature, bubbles per minute, and timing with a digital stop clock will be taken as part of the experiment. In the study, we grew barley and corn in elevated or low CO2 (370, 700 ppm) and optimal soil N and discovered that high CO2 during photosynthesis has negative effects on the rubisco activase and glutathione peroxidase (gpse) molecule. The thermal optimum of a PSII experiment is similar to that of a C3 or C4 plants electron transfer chain. The summary diagram (Figure 15) classifies the effects of temperature on photosynthesis according to the speed of temperature change and the extent of its variation. The ideal temperature for photosynthesis is between 70-85 degrees Fahrenheit. Developmental Plasticity at High Temperature. The formation of reducing power and the synthesis of ATP have a thermal sensitivity close to that of electron transfer. Figure 4 shows CO2 assimilation in pea grown at 10 or 25C. In most cases its response to temperature is rapidly reversible between about 10 and 34C. What Is Rubisco Role In Photosynthesis Crash Course, The Process Of Photosynthesis: How Plants Create The Energy We Need To Live, The Potential Dangers Of Overdosing On Neem: What You Need To Know. Most plants, in general, respond well to normal temperature fluctuations. Plant Physiol.181, 399-411. Plant Physiol. Photosynthesis is, for lack of a better explanation, how plants and some other organisms "eat." Because photosynthesis is a chemical reaction, most of the chemical reactions are accelerated as the temperature rises. To help the organism remain functional, we hope to be able to generate enough primary products. (2014). The reason for this is that the enzymes that are responsible for photosynthesis are less active at lower temperatures. [5] Plants from the same individual by vegetative reproduction. Performance cookies are used to understand and analyze the key performance indexes of the website which helps in delivering a better user experience for the visitors. Black line is linear regression line, and grey line is 1:1 line. It has six letters (N.6A 5B7, N.M.H., N.P.A.H.). As a result, the oxygenase function of the Rubisco was released, resulting in the loss of fixed carbon. Its content can reach 800 to 2000 ppm depending on the plant in C4: that is to say contents from 2 to 5 times higher than its current atmospheric content. At high temperatures, enzymes are denatured. Effect of temperature on net CO2 uptake, stomatal conductance for CO2 and quantum yield of photosystem II photochemistry of dehydrated pea leaves. So, while extremely high temperatures can have a negative impact on photosynthesis, mild increases in temperature can actually improve the rate at which this process occurs. Plant Physiol.182, 1404-10419. These cookies ensure basic functionalities and security features of the website, anonymously. 182, 16241635, Steponkus PL & Lanphear FO (1968). It shows minimum, optimum and maximum temperatures. The presence of these agents influences the functions of photosynthetic carbon assimilation enzymes, the photoactive opening of stomata, metabolite accumulation, and pigment composition, among other things [1, 2]. Plants in cold environments or with a cold growing season have a higher photosynthesis at low temperatures. Oxygen is a gas that is used by animals to breathe. What is the optimal conditions for photosynthesis? The Power Of Neem DE: A Natural And Effective Insecticide, Getting Rid Of Pesky Caterpillars With Neem Oil, Understanding The Benefits And Risks Of Giving Neem To Your Dog. When photosynthesis is at its optimum, the temperature ranges from 35C to 40C. What is optimum pH and temperature in enzymes? Because temperature increases, the number of collisions increases, resulting in a faster rate of photosynthesis. This is hardening. In a process driven by light energy, glucose molecules (or other sugars) are constructed from water and carbon dioxide, and oxygen is released as a byproduct. When the ambient airs O2 content is low (1% for example), or when CO2 levels are high (for example, higher), CO2 occupies a high number of active sites on the Rubisco. Finally, low temperatures can also affect the amount of light that is able to reach the leaves, which can reduce the amount of photosynthesis that can take place. The Potential Risks And Benefits Of Neem Oil Soap: Is It Safe To Use? Hypothermia, if left untreated, can cause irreversible damage to your heart and respiratory system. The optimal temperature is between 10-20 degrees C (50-68 degrees F), though this can vary widely depending on the plant type. The chemistry of photosynthesis, whether it takes place in water or air, remains constant. The articles in the Encyclopedia of the Environment are made available under the terms of the Creative Commons BY-NC-SA license, which authorizes reproduction subject to: citing the source, not making commercial use of them, sharing identical initial conditions, reproducing at each reuse or distribution the mention of this Creative Commons BY-NC-SA license. The evaporative power of the air is also regulated in most cases during the measurements. As in any other enzyme-controlled reaction, the rate at which photosynthesis takes place is affected by temperature. The optimum air temperature for photosynthesis is one of the major parameters in data-driven and process-based photosynthesis models that estimate the gross primary production (GPP) of vegetation under a changing climate. 1). When the temperature decreases, the photosynthesis process also decreases. The process is truly dependent on three main factors. When the temperature rises too high, the enzymes that aid in catalyzing the reaction begin to denature, rendering them ineffective. The optimal temperature for photosynthesis at its most efficient is 35 to 40 degrees Celsius. USA 97, 13430-13435, Falk S, Maxwell DP, Laudenbach DE & Huner NPA (1996). Your plants can grow in healthy and strong environments if you use the right tools. The optimum temperature varies depending on the type of plant, but is typically between 20 and 30 degrees Celsius. However, when the temperature reaches more than 40 degrees Celsius, photosynthesis slows down. Grow lights emit artificial light that allows farmers and growers to simulate photosynthesis. How is the carbon from carbon dioxide CO2 present in the atmosphere integrated.

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what is optimum temperature for photosynthesis